As a member of Dr. Nicolas Fawzi’s laboratory at Brown University, Ryan Puterbaugh focused his research on understanding the molecular mechanisms that govern the behavior of the disordered TDP-43 prion-like domain. On August 4, 2026, Ryan successfully defended his PhD thesis entitled “Understanding the Biomolecular Interactions of the Disordered TDP-43 Prion-Like Domain in the Context of both Health and Disease” in Room 107 at 70 Ship Street.

Following the defense, Ryan celebrated this significant milestone with his advisor, Dr. Nicolas Fawzi, as well as Brown students, faculty, family, and friends.

Congratulations, Dr. Puterbaugh!